| SETRES SETRES
is major collaborative research effort to
expand our fundamental understanding of
relationships among forest productivity,
genetics, and resource availability using
loblolly pine as a model species.
Collaborators include numerous scientists and
graduate students from N.C. State University,
the U.S. Forest Services
Biological Foundations of Southern Forest
Productivity and Sustainability Work Unit at
Research Triangle Park, Duke University,
and Virginia Tech. The first SETRES study was
established in 1992 and includes a 2 x 2
factorial combination of nutrient (no
addition and optimum nutrition) and water (no
addition and well watered) treatments imposed
on large plots within what was then a 7-year
loblolly pine stand. The second study,
established in 1995, includes five families
from each of two extreme loblolly pine
provenances (Atlantic Coastal Plain and Lost
Pines of Texas provenances) grown under low
(no nutrients added) and optimum (nutrients
added every year) nutrition regimes. Numerous
assessments have been made and are continuing
on these two studies including
photosynthesis, tissue growth and maintenance
respiration, biomass production, leaf area,
carbon allocation, soil respiration, net
ecosystem productivity, tissue nutrient
concentrations, soil carbon pools and fluxes,
soil nutrient pools and fluxes, litterfall,
forest floor development and decomposition,
soil N mineralization, soil microbial
communities, phenology of above and
below-ground growth, wood quality, stem
quality, crown development, intra-specific
competition and stand development, tree water
relations, hydrologic budgets, microclimatic
conditions, and spectral reflectance to
determine leaf area and foliar nutrition. The
assessments are being integrated through the
use of several physiologically based
production models and have allowed us to
scale from the tissue to the landscape level.
To-date, over twenty graduate students
have worked on thesis projects at the site
and almost 70 papers
and theses have been published. During
the next few years, analysis and modeling
efforts will focus on better understanding
the long term of effects of elevated levels
of resource availability on ecosystem
productivity, nutrient cycling,
intra-specific competition, and carrying
capacity relationships.
Key results to date include:
- Nutrient additions have increase
biomass production by 2.5x this
dry sand hill site.
Irrigation has had little impact on
production.
- Gains in production with increased
nutrient availability have resulted
from increases in leaf area and
growth efficiency.
- Stemwood growth efficiency gains
resulted from a strong shift in
biomass allocation from fine roots to
stemwood. On control plots fine roots
accounted for 20% of production and
only 8% on fertilized plots although
the absolute production of fine roots
remained nearly the same for the two
treatments.
- Nutrient additions not only increased
carbon gain but also dramatically
reduced the period of time needed for
site to regain positive net ecosystem
productivity.
- Positive carbon gain responses of
individual tress to elevated levels
of C02 were limited by low nutrient
availability.
- Increasing nutrient availability
reduced within stand heterogeneity in
tree size leading to slower stand
differentiation into crown classes.
This increased uniformity can be
expected to impact future stand
development, structure, and yield in
ways for which current growth and
yield models do not account.
- Ecosystem retention of added nitrogen
was very high.
- Stand leaf area was easily and
accurately estimated using Landsat
imagery.
- Provenance and family effects have
been highly significant but small
compared to the nutrient effects.
- Few traits have shown any genetic x
environmental interaction even though
the range in genetic material and
nutrient treatments represented
extreme differences.
In addition to serving as a base for
considerable research, these studies also
serve as very important education and
demonstration sites. Over the past 15 years,
the studies have been visited by several
thousand forest managers, scientists,
undergraduates, and graduate students from
around the world.
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